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Why does unique_ptr take two template parameters when shared_ptr only takes one?

Both unique_ptr and shared_ptr accept a custom destructor to call on the object they own. But in the case of unique_ptr, the destructor is passed as a template parameter of the class, wherease the type of shared_ptr's custom destructor is to be specified as a template parameter of the constructor.

template <class T, class D = default_delete<T>>
class unique_ptr
{
    unique_ptr(T*, D&); //simplified
    ...
};

and

template<class T>
class shared_ptr
{
    template<typename D>
    shared_ptr(T*, D); //simplified
    ...
};

I can’t see why such difference. What requires that?